Silurian Period

The Silurian is a major division of the geologic timescale that started about 443.8 ± 1.5 Ma. During the Silurian, Gondwana continued a slow southward drift to high southern latitudes, but there is evidence that the Silurian ice caps were less extensive than those of the late Ordovician glaciation. The melting of ice caps and glaciers contributed to a rise in sea levels, recognizable from the fact that Silurian sediments overlie eroded Ordovician sediments, forming an unconformity. Other cratons and continent fragments drifted together near the equator, starting the formation of a second supercontinent known as Euramerica. The vast ocean of Panthalassa covered most of the northern hemisphere. Other minor oceans include Proto-Tethys, Paleo-Tethys, Rheic Ocean, a seaway of Iapetus Ocean (now in between Avalonia and Laurentia), and newly formed Ural Ocean.

Devonian Period

The Devonian spanned roughly from 419 to 359 Ma. The period was a time of great tectonic activity, as Laurasia and Gondwana drew closer together. The continent Euramerica (or Laurussia) was created in the early Devonian by the collision of Laurentia and Baltica, which rotated into the natural dry zone along the Tropic of Capricorn. In these near-deserts, the Old Red Sandstone sedimentary beds formed, made red by the oxidized iron (hematite) characteristic of drought conditions. Near the equator Pangaea began to consolidate from the plates containing North America and Europe, further raising the northern Appalachian Mountains and forming the Caledonian Mountains in Great Britain and Scandinavia. The southern continents remained tied together in the supercontinent of Gondwana. The remainder of modern Eurasia lay in the Northern Hemisphere. Sea levels were high worldwide, and much of the land lay submerged under shallow seas. The deep, enormous Panthalassa (the "universal ocean") covered the rest of the planet. Other minor oceans were Paleo-Tethys, Proto-Tethys, Rheic Ocean and Ural Ocean (which was closed during the collision with Siberia and Baltica).

Carboniferous Period

The Carboniferous extends from about 358.9 ± 0.4 to about 298.9 ± 0.15 Ma.

A global drop in sea level at the end of the Devonian reversed early in the Carboniferous; this created the widespread epicontinental seas and carbonate deposition of the Mississippian. There was also a drop in south polar temperatures; southern Gondwana was glaciated throughout the period, though it is uncertain if the ice sheets were a holdover from the Devonian or not. These conditions apparently had little effect in the deep tropics, where lush coal swamps flourished within 30 degrees of the northernmost glaciers. A mid-Carboniferous drop in sea-level precipitated a major marine extinction, one that hit crinoids and ammonites especially hard. This sea-level drop and the associated unconformity in North America separate the Mississippian Period from the Pennsylvanian period.

The Carboniferous was a time of active mountain building, as the supercontinent Pangea came together. The southern continents remained tied together in the supercontinent Gondwana, which collided with North America-Europe (Laurussia) along the present line of eastern North America. This continental collision resulted in the Hercynian orogeny in Europe, and the Alleghenian orogeny in North America; it also extended the newly uplifted Appalachians southwestward as the Ouachita Mountains. In the same time frame, much of present eastern Eurasian plate welded itself to Europe along the line of the Ural mountains. There were two major oceans in the Carboniferous the Panthalassa and Paleo-Tethys. Other minor oceans were shrinking and eventually closed the Rheic Ocean (closed by the assembly of South and North America), the small, shallow Ural Ocean (which was closed by the collision of Baltica, and Siberia continents, creating the Ural Mountains) and Proto-Tethys Ocean.